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FINDING CURES TOGETHER<sup>SM</sup>

Fellows of the AACR Academy

Jerry Adams, PhD

Jerry Adams, PhD
Walter & Eliza Hall Institute of Medical Research, Melbourne, Australia

Bruce M. Alberts, PhD

Bruce M. Alberts, PhD
University of California, San Francisco, San Francisco, California

James P. Allison, PhD

James P. Allison, PhD
University of Texas MD Anderson Cancer Center, Houston, Texas

Kenneth C. Anderson, MD

Kenneth C. Anderson, MD
Dana-Farber Cancer Institute, Harvard Medical School, Boston, Massachusetts

Karen H. Antman, MD

Karen H. Antman, MD
Boston University School of Medicine, Boston, Massachusetts

Frederick R. Appelbaum, MD

Frederick R. Appelbaum, MD
Fred Hutchinson Cancer Research Center, Seattle, Washington

Carlos L. Arteaga, MD

Carlos L. Arteaga, MD
Vanderbilt University, Nashville, Tennessee

Alan Ashworth, PhD, FRS

Alan Ashworth, PhD, FRS
UCSF Helen Diller Family Comprehensive Cancer Center; University of California, San Francisco, San Francisco, California

David Baltimore, PhD

David Baltimore, PhD
California Institute of Technology, Pasadena, California

Mariano Barbacid, PhD

Mariano Barbacid, PhD
Centro Nacional de Investigaciones Oncológicas, Madrid, Spain

Françoise Barré-Sinoussi, PhD

Françoise Barré-Sinoussi, PhD
Institut Pasteur, Paris, France

Dafna Bar-Sagi, PhD

Dafna Bar-Sagi, PhD
NYU Langone Health, New York

José Baselga, MD, PhD

José Baselga, MD, PhD
Memorial Sloan Kettering Cancer Center, New York, New York

Stephen B. Baylin, MD

Stephen B. Baylin, MD
Sidney Kimmel Comprehensive Cancer Center; Johns Hopkins University School of Medicine, Baltimore, Maryland

Paul Berg, PhD

Paul Berg, PhD
Stanford University School of Medicine, Stanford, California

René Bernards, PhD

René Bernards, PhD
The Netherlands Cancer Institute, Amsterdam, Netherlands

Anton Berns, PhD

Anton Berns, PhD
The Netherlands Cancer Institute, Amsterdam, Netherlands

Joseph R. Bertino, MD

Joseph R. Bertino, MD
The Cancer Institute of New Jersey, New Brunswick, New Jersey

Bruce A. Beutler, MD

Bruce A. Beutler, MD
Center for the Genetics of Host Defense; University of Texas Southwestern Medical Center, Dallas, Texas

Sir Adrian P. Bird, PhD

Sir Adrian P. Bird, PhD
University of Edinburgh, Scotland

J. Michael Bishop, MD

J. Michael Bishop, MD
University of California, San Francisco, San Francisco, California

Mina J. Bissell, PhD

Mina J. Bissell, PhD
Lawrence Berkeley National Laboratory, Berkeley, California

Elizabeth H. Blackburn, PhD

Elizabeth H. Blackburn, PhD
The Salk Institute, La Jolla, California

Günter Blobel, MD, PhD

Günter Blobel, MD, PhD
deceased (1936-2018)

Clara D. Bloomfield, MD

Clara D. Bloomfield, MD
The Ohio State University Comprehensive Cancer Center, James Cancer Hospital and Solove Research Institute, Columbus, Ohio

David Botstein, PhD

David Botstein, PhD
Princeton University, Princeton, New Jersey

Sydney Brenner, MBBCh, DPhil

Sydney Brenner, MBBCh, DPhil
deceased (1927-2019)

Angela H. Brodie, PhD

Angela H. Brodie, PhD
deceased (1934-2017)

Joan S. Brugge, PhD

Joan S. Brugge, PhD
Ludwig Center; Harvard Medical School, Boston, Massachusetts

Michael A. Caligiuri, MD

Michael A. Caligiuri, MD
The City of Hope National Medical Center, Duarte, California

Lewis C. Cantley, PhD

Lewis C. Cantley, PhD
Meyer Cancer Center, Weill Cornell Medical College, New York, New York

Mario R. Capecchi, PhD

Mario R. Capecchi, PhD
University of Utah School of Medicine, Salt Lake City, Utah

Webster K. Cavenee, PhD

Webster K. Cavenee, PhD
Ludwig Institute for Cancer Research; University of California, San Diego, San Diego, California

Thomas R. Cech, PhD

Thomas R. Cech, PhD
BioFrontiers Institute, University of Colorado, Boulder, Colorado

Bruce A. Chabner, MD

Bruce A. Chabner, MD
Massachusetts General Hospital Cancer Center, Boston, Massachusetts

Martin Chalfie, PhD

Martin Chalfie, PhD
Columbia University, New York, New York

Pierre Chambon, MD

Pierre Chambon, MD
University of Strasbourg Institute for Advanced Study, Strasburg, France

Emmanuelle Charpentier, PhD

Emmanuelle Charpentier, PhD
Max Planck Institute for Infection Biology, Berlin, Germany

Zhu Chen, MD, PhD

Zhu Chen, MD, PhD
12th Standing Committee of the National People’s Congress; Chinese Medical Association, Beijing, China

Aaron J. Ciechanover, MD, DSc

Aaron J. Ciechanover, MD, DSc
Technion - Israel Institute of Technology, Haifa, Israel

Bayard D. Clarkson, MD

Bayard D. Clarkson, MD
Memorial Sloan Kettering Cancer Center, New York, New York

Hans Clevers, MD, PhD

Hans Clevers, MD, PhD
Royal Netherlands Academy of Arts and Sciences, Hubrecht Institute, Utrecht, The Netherlands

Donald S. Coffey, PhD

Donald S. Coffey, PhD
deceased (1932-2017)

Stanley N. Cohen, MD

Stanley N. Cohen, MD
Stanford University, Stanford, California

Suzanne Cory, PhD

Suzanne Cory, PhD
Walter and Eliza Hall Institute of Medical Research; University of Melbourne, Melbourne, Australia

Lisa M. Coussens, PhD

Lisa M. Coussens, PhD
Knight Cancer Institute; Oregon Health & Science University, Portland, Oregon

Carlo M. Croce, MD

Carlo M. Croce, MD
Institute of Genetics, The Ohio State University School of Medicine, Columbus, Ohio

Tom Curran, PhD

Tom Curran, PhD
Children's Mercy Hospital, Kansas City, Missouri

George Q. Daley, MD, PhD

George Q. Daley, MD, PhD
Harvard Medical School, Cambridge, Massachusetts

Riccardo Dalla-Favera, MD

Riccardo Dalla-Favera, MD
Institute for Cancer Genetics, Columbia University, New York, New York

Chi Van Dang, MD, PhD

Chi Van Dang, MD, PhD
Ludwig Institute for Cancer Research, New York, New York

James E. Darnell Jr., MD

James E. Darnell Jr., MD
The Rockefeller University, New York, New York

Nancy E. Davidson, MD

Nancy E. Davidson, MD
Fred Hutchinson Cancer Research Center; Seattle Cancer Care Alliance; University of Washington, Seattle, Washington

Titia de Lange, PhD

Titia de Lange, PhD
Anderson Center for Cancer Research; Rockefeller University, New York, New York

Ronald A. DePinho, MD

Ronald A. DePinho, MD
University of Texas MD Anderson Cancer Center, Houston, Texas

Susan D. Desmond-Hellman, MD, MPH

Susan D. Desmond-Hellman, MD, MPH
Bill & Melinda Gates Foundation, Seattle, Washington; University of California, San Francisco, San Francisco, California

Vincent T. DeVita Jr., MD

Vincent T. DeVita Jr., MD
Yale University School of Medicine, New Haven, Connecticut

John E. Dick, PhD

John E. Dick, PhD
Ontario Institute for Cancer Research, Toronto, Ontario

Vishva M. Dixit, MD

Vishva M. Dixit, MD
Genentech, Inc., San Francisco, California

Jennifer A. Doudna, PhD

Jennifer A. Doudna, PhD
University of California, Berkeley, Berkeley, California

Brian J. Druker, MD

Brian J. Druker, MD
Oregon Health & Science University, Portland, Oregon

Raymond N. DuBois, MD, PhD

Raymond N. DuBois, MD, PhD
Medical University of South Carolina, Charleston, South Carolina

Lawrence H. Einhorn, MD

Lawrence H. Einhorn, MD
Indiana University School of Medicine, Indianapolis, Indiana

Robert N. Eisenman, PhD

Robert N. Eisenman, PhD
Fred Hutchinson Cancer Research Center, Seattle, Washington

Stephen J. Elledge, PhD

Stephen J. Elledge, PhD
Harvard Medical School, Boston, Massachusetts

Gerard I. Evan, PhD, FRS

Gerard I. Evan, PhD, FRS
University of Cambridge, Cambridge, United Kingdom

Sir Martin J. Evans, PhD

Sir Martin J. Evans, PhD
Cardiff University, Cardiff, United Kingdom

Ronald M. Evans, PhD

Ronald M. Evans, PhD
Gene Expression Laboratory; Salk Institute for Biological Studies, La Jolla, California

Emmanuel Farber, MD, PhD

Emmanuel Farber, MD, PhD
deceased (1918-2014)

Napoleone Ferrara, MD

Napoleone Ferrara, MD
Moores Cancer Center; University of California San Diego School of Medicine, San Diego, California

Isaiah J. Fidler, DVM, PhD

Isaiah J. Fidler, DVM, PhD
University of Texas MD Anderson Cancer Center, Houston, Texas

Andrew Z. Fire, PhD

Andrew Z. Fire, PhD
Stanford University School of Medicine, Stanford, California

Bernard Fisher, MD

Bernard Fisher, MD
deceased (1918-2019)

Joseph F. Fraumeni Jr., MD

Joseph F. Fraumeni Jr., MD
National Cancer Institute, National Institutes of Health, Bethesda, Maryland

Gordon J. Freeman, PhD

Gordon J. Freeman, PhD
Harvard Medical School, Boston, Massachusettes

Emil Frei III, MD

Emil Frei III, MD
deceased (1924-2013)

Emil J Freireich, MD

Emil J Freireich, MD
University of Texas MD Anderson Cancer Center, Houston, Texas

Elaine Fuchs, PhD

Elaine Fuchs, PhD
Rockefeller University, New York, New York

Robert C. Gallo, MD

Robert C. Gallo, MD
University of Maryland School of Medicine, Baltimore, Maryland

Judy E. Garber, MD, MPH

Judy E. Garber, MD, MPH
Center for Cancer Genetics and Prevention, Dana-Farber Cancer Institute, Boston, Massachusetts

Levi A. Garraway, MD, PhD,

Levi A. Garraway, MD, PhD,
Eli Lilly and Company, Indianapolis, Indiana

Walter Gilbert, PhD

Walter Gilbert, PhD
BioVentures Investors, Wellesley Hills, Massachusetts

Dwight Gary Gilliland, MD, PhD

Dwight Gary Gilliland, MD, PhD
Fred Hutchinson Cancer Research Center, Seattle, Washington

Alfred G. Gilman, MD, PhD

Alfred G. Gilman, MD, PhD
deceased (1941-2015)

Laurie H. Glimcher, MD

Laurie H. Glimcher, MD
Dana-Farber Cancer Institute, Harvard Medical School, Boston, Massachusetts

Joe W. Gray, PhD

Joe W. Gray, PhD
Oregon Health & Science University, Portland, Oregon

Sir Mel F. Greaves, PhD

Sir Mel F. Greaves, PhD
The Institute of Cancer Research, London, United Kingdom

Philip D. Greenberg, MD

Philip D. Greenberg, MD
Fred Hutchinson Cancer Research Center, Seattle, Washington

Carol W. Greider, PhD

Carol W. Greider, PhD
Johns Hopkins University School of Medicine, Baltimore, Maryland

Roger C.L. Guillemin, MD, PhD

Roger C.L. Guillemin, MD, PhD
The Salk Institute for Biological Studies, La Jolla, California

Sir John B. Gurdon, DPhil

Sir John B. Gurdon, DPhil
Wellcome Trust; Cancer Research UK Gurdon Institute, Cambridge, United Kingdom

Daniel A. Haber, MD, PhD

Daniel A. Haber, MD, PhD
Massachusetts General Hospital Cancer Center; Harvard Medical School, Boston, Massachusetts

William N. Hait, MD, PhD

William N. Hait, MD, PhD
Janssen Research & Development, Raritan, New Jersey

Douglas Hanahan, PhD

Douglas Hanahan, PhD
Swiss Institute for Experimental Cancer Research, Lausanne, Switzerland

Leland H. Hartwell, PhD

Leland H. Hartwell, PhD
Arizona State University Biodesign Institute, Tempe, Arizona

Avram Hersko, MD, PhD

Avram Hersko, MD, PhD
Technion-Israel Institute of Technology, Haifa, Israel; New York University School of Medicine, New York, New York

Jules A. Hoffmann, PhD

Jules A. Hoffmann, PhD
University of Strasbourg Institute for Advanced Study, Strasbourg, France

James F. Holland, MD

James F. Holland, MD
deceased (1925-2018)

Jimmie C. Holland, MD

Jimmie C. Holland, MD
deceased (1928-2017)

Waun Ki Hong, MD

Waun Ki Hong, MD
deceased (1942-2019)

Tasuku Honjo, MD, PhD

Tasuku Honjo, MD, PhD
Kyoto University Institute for Advanced Study, Kyoto, Japan

Leroy E. Hood, MD, PhD

Leroy E. Hood, MD, PhD
Institute for Systems Biology, Seattle, Washington

H. Robert Horvitz, PhD

H. Robert Horvitz, PhD
Massachusetts Institute of Technology, Cambridge, Massachusetts

Susan B. Horwitz, PhD

Susan B. Horwitz, PhD
Albert Einstein College of Medicine, New York, New York

Sir R. Timothy Hunt, PhD

Sir R. Timothy Hunt, PhD
Formerly affiliated with Cancer Research UK, London, United Kingdom

Tony Hunter, PhD

Tony Hunter, PhD
The Salk Institute, La Jolla, California

Richard O. Hynes, PhD

Richard O. Hynes, PhD
Massachusetts Institute of Technology, Cambridge, Massachusetts

Tyler Jacks, PhD

Tyler Jacks, PhD
David H. Koch Institute for Integrative Cancer Research; Massachusetts Institute of Technology, Cambridge, Massachusetts

Rudolf Jaenisch, MD

Rudolf Jaenisch, MD
Whitehead Institute for Biomedical Research; Massachusetts Institute of Technology, Cambridge, Massachusetts

Elizabeth M. Jaffee, MD

Elizabeth M. Jaffee, MD
Sidney Kimmel Comprehensive Cancer Center, The Johns Hopkins University School of Medicine, Baltimore, Maryland

Peter A. Jones, PhD, DSc

Peter A. Jones, PhD, DSc
Van Andel Research Institute, Grand Rapids, Michigan

V. Craig Jordan, CMG, OBE, PhD, DSc, FMedSci

V. Craig Jordan, CMG, OBE, PhD, DSc, FMedSci
University of Texas MD Anderson Cancer Center, Houston, Texas

Carl H. June, MD

Carl H. June, MD
University of Pennsylvania, Philadelphia, Pennsylvania

William G. Kaelin Jr., MD

William G. Kaelin Jr., MD
Dana-Farber Cancer Institute, Harvard Medical School, Boston, Massachusetts

Yuet Wai Kan, MD

Yuet Wai Kan, MD
University of California, San Francisco, San Francisco, California

Michael Karin, PhD

Michael Karin, PhD
University of California, San Diego, La Jolla, California

Michael B. Kastan, MD, PhD

Michael B. Kastan, MD, PhD
Duke Cancer Institute; Duke University School of Medicine, Durham, NC

Mary-Claire King, PhD

Mary-Claire King, PhD
University of Washington, Seattle, Washington

Kenneth W. Kinzler, PhD

Kenneth W. Kinzler, PhD
Johns Hopkins Kimmel Cancer Center, Baltimore, Maryland

Richard D. Klausner, MD

Richard D. Klausner, MD
Juno Therapeutics, Seattle, Washington

Eva Klein, MD, PhD

Eva Klein, MD, PhD
Karolinska Institute, Stockholm, Sweden

George Klein, MD, PhD

George Klein, MD, PhD
deceased (1925-2016)

Alfred G. Knudson Jr., MD, PhD

Alfred G. Knudson Jr., MD, PhD
deceased (1922-2016)

Brian K. Kobilka, MD

Brian K. Kobilka, MD
Stanford University School of Medicine, Stanford, California

Richard D. Kolodner, PhD

Richard D. Kolodner, PhD
Ludwig Institute for Cancer Research; University of California San Diego School of Medicine, San Diego, California

Roger D. Kornberg, PhD

Roger D. Kornberg, PhD
Stanford University School of Medicine, Stanford, California

Margaret L. Kripke, PhD

Margaret L. Kripke, PhD
University of Texas MD Anderson Cancer Center, Houston, Texas

Eric S. Lander, PhD

Eric S. Lander, PhD
The Broad Institute of MIT and Harvard, Cambridge, Massachusetts

Sir David P. Lane, PhD

Sir David P. Lane, PhD
Technology and Research (A*STAR), Singapore; Ludwig Institute for Cancer Research, New York, New York

Philip Leder, MD

Philip Leder, MD
Harvard Medical School, Boston, Massachusetts

Robert J. Lefkowitz, MD

Robert J. Lefkowitz, MD
Duke University Medical Center, Durham, North Carolina

Arnold J. Levine, PhD

Arnold J. Levine, PhD
Institute for Advanced Study, Princeton, New Jersey

Arthur D. Levinson, PhD

Arthur D. Levinson, PhD
Calico Life Sciences LLC, San Francisco, California

Ronald Levy, MD

Ronald Levy, MD
Stanford University School of Medicine, Palo Alto, California

Frederick P. Li, MD

Frederick P. Li, MD
deceased (1940-2015)

Tomas Lindahl, MD, FRS

Tomas Lindahl, MD, FRS
Francis Crick Institute, London, United Kingdom

David M. Livingston, MD

David M. Livingston, MD
Dana-Farber Cancer Institute, Harvard Medical School, Boston, Massachusetts

Lawrence A. Loeb, MD, PhD

Lawrence A. Loeb, MD, PhD
University of Washington School of Medicine, Seattle, Washington

Scott W. Lowe, PhD

Scott W. Lowe, PhD
Memorial Sloan Kettering Cancer Center, New York, New York

Douglas R. Lowy, MD

Douglas R. Lowy, MD
National Cancer Institute, Bethesda, Maryland

Henry T. Lynch, MD

Henry T. Lynch, MD
deceased (1928-2019)

Tak W. Mak, PhD

Tak W. Mak, PhD
Princess Margaret Cancer Centre; University of Toronto, Toronto, Ontario

Elaine R. Mardis, PhD

Elaine R. Mardis, PhD
Institute for Genomic Medicine, Nationwide Children’s Hospital, Columbus, Ohio

Paul A. Marks, MD

Paul A. Marks, MD
Memorial Sloan Kettering Cancer Center, New York, New York

Joan Massagué, PhD

Joan Massagué, PhD
Memorial Sloan Kettering Cancer Center, New York, New York

Lynn M. Matrisian, PhD

Lynn M. Matrisian, PhD
Pancreatic Cancer Action Network, Manhattan Beach, California

Frank McCormick, PhD

Frank McCormick, PhD
University of California, San Francisco, San Francisco, California

John Mendelsohn, MD

John Mendelsohn, MD
deceased (1936-2019)

Donald Metcalf, MD

Donald Metcalf, MD
deceased (1929-2014)

Enrico Mihich, MD

Enrico Mihich, MD
deceased (1928-2016)

Beatrice Mintz, PhD

Beatrice Mintz, PhD
Fox Chase Cancer Center, Philadelphia, Pennsylvania

Paul L. Modrich, PhD

Paul L. Modrich, PhD
Duke University Medical Center, Durham, North Carolina

Luc Montagnier, MD

Luc Montagnier, MD
Pasteur Institute, Paris, France

Harold L. Moses, MD

Harold L. Moses, MD
Vanderbilt-Ingram Comprehensive Cancer Center, Nashville, Tennessee

Larry Norton, MD

Larry Norton, MD
Memorial Sloan Kettering Cancer Center, New York, New York

Peter C. Nowell, MD

Peter C. Nowell, MD
deceased (1928-2016)

Sir Paul M. Nurse, PhD

Sir Paul M. Nurse, PhD
Francis Crick Institute, London, United Kingdom

Christiane Nüsslein-Volhard, PhD

Christiane Nüsslein-Volhard, PhD
Max Planck Institute for Developmental Biology, Tübingen, Germany

Olufunmilayo I. Olopade, MD

Olufunmilayo I. Olopade, MD
University of Chicago Pritzker School of Medicine, Chicago, Illinois

Moshe Oren, PhD

Moshe Oren, PhD
Moross Integrated Cancer Center; Weizmann Institute of Science, Rehovot, Israel

Arthur B. Pardee, PhD

Arthur B. Pardee, PhD
deceased (1921-2019)

Sir Richard Peto, FRS

Sir Richard Peto, FRS
University of Oxford, Oxford, United Kingdom

Martine J. Piccart, MD, PhD

Martine J. Piccart, MD, PhD
Jules Bordet Institute, Brussels, Belgium

Sir Bruce A.J. Ponder, PhD

Sir Bruce A.J. Ponder, PhD
Cancer Research UK Cambridge Research Institute; University of Cambridge, Cambridge, United Kingdom

Carol L. Prives, PhD

Carol L. Prives, PhD
Columbia University, New York, New York

Sir Richard J. Roberts, PhD

Sir Richard J. Roberts, PhD
New England Biolabs, Inc., Ipswich, Massachusetts

Irwin A. Rose, PhD

Irwin A. Rose, PhD
deceased (1926-2015)

Steven A. Rosenberg, MD, PhD

Steven A. Rosenberg, MD, PhD
National Cancer Institute, Bethesda, Maryland

Janet D. Rowley, MD

Janet D. Rowley, MD
deceased (1925 - 2013)

Frederick Sanger, PhD

Frederick Sanger, PhD
deceased (1918 - 2013)

Alan C. Sartorelli, PhD

Alan C. Sartorelli, PhD
deceased (1931-2015)

Charles L. Sawyers, MD

Charles L. Sawyers, MD
Memorial Sloan Kettering Cancer Center, New York, New York

Andrew V. Schally, PhD

Andrew V. Schally, PhD
University of Miami, Miami, Florida

Joseph Schlessinger, PhD

Joseph Schlessinger, PhD
Yale School of Medicine, New Haven, Connecticut

Robert D. Schreiber, PhD

Robert D. Schreiber, PhD
Bursky Center for Human Immunology and Immunotherapy Programs, Washington University School of Medicine, St. Louis, Missouri

Phillip A. Sharp, PhD

Phillip A. Sharp, PhD
Massachusetts Institute of Technology, Cambridge, Massachusetts

Arlene H. Sharpe, MD, PhD

Arlene H. Sharpe, MD, PhD
Harvard Medical School, Boston, Massachusetts

Norman E. Sharpless, MD

Norman E. Sharpless, MD
Food and Drug Administration, Silver Spring, Maryland

Charles J. Sherr, MD, PhD

Charles J. Sherr, MD, PhD
St. Jude Children’s Research Hospital, Memphis, Tennessee

Osamu Shimomura, PhD

Osamu Shimomura, PhD
deceased (1928-2018)

Dennis J. Slamon, MD, PhD

Dennis J. Slamon, MD, PhD
University of California, Los Angeles, Los Angeles, California

Oliver Smithies, DPhil

Oliver Smithies, DPhil
deceased (1925-2017)

Michael B. Sporn, MD

Michael B. Sporn, MD
Geisel School of Medicine at Dartmouth, Hanover, New Hampshire

Louis M. Staudt, MD, PhD

Louis M. Staudt, MD, PhD
National Cancer Institute, Bethesda, Maryland

Bruce W. Stillman, PhD

Bruce W. Stillman, PhD
Cold Spring Harbor Laboratory, Cold Spring Harbor, New York

Sir Michael R. Stratton, MBBS, PhD

Sir Michael R. Stratton, MBBS, PhD
Wellcome Trust Sanger Institute, Cambridge, United Kingdom

Louise C. Strong, MD

Louise C. Strong, MD
University of Texas MD Anderson Cancer Center, Houston, Texas

Takashi Sugimura, MD

Takashi Sugimura, MD
Toho University; National Cancer Center, Tokyo, Japan

Sir John E. Sulston, PhD

Sir John E. Sulston, PhD
deceased (1942-2018)

Jack W. Szostak, PhD

Jack W. Szostak, PhD
Massachusetts General Hospital, Boston, Massachusetts

Craig B. Thompson, MD

Craig B. Thompson, MD
Memorial Sloan Kettering Cancer Center, New York, New York

Roger Y. Tsien, PhD

Roger Y. Tsien, PhD
deceased (1952-2016)

Axel Ullrich, PhD

Axel Ullrich, PhD
Max Planck Institute of Biochemistry, Munich, Germany

Arthur C. Upton, MD

Arthur C. Upton, MD
deceased (1923-2015)

George F. Vande Woude, PhD

George F. Vande Woude, PhD
Van Andel Research Institute, Grand Rapids, Michigan

Inder M. Verma, PhD

Inder M. Verma, PhD
Formerly affiliated with the Salk Institute for Biological Sciences, La Jolla, California

Bert Vogelstein, MD

Bert Vogelstein, MD
Ludwig Center for Cancer Genetics and Therapeutics, Johns Hopkins University School of Medicine, Baltimore, Maryland

Peter K. Vogt, PhD

Peter K. Vogt, PhD
The Scripps Research Institute, La Jolla, California

Daniel D. Von Hoff, MD

Daniel D. Von Hoff, MD
Translational Genomics Research Institute, Phoenix, Arizona

Karen H. Vousden, PhD

Karen H. Vousden, PhD
Cancer Research UK, London, United Kingdom

Geoffrey M. Wahl, PhD

Geoffrey M. Wahl, PhD
The Salk Institute for Biological Studies, La Jolla, California

James D. Watson, PhD

James D. Watson, PhD
Cold Spring Harbor Laboratory, Cold Spring Harbor, New York

Lee W. Wattenberg, MD

Lee W. Wattenberg, MD
deceased (1921-2014)

Robert A. Weinberg, PhD

Robert A. Weinberg, PhD
Ludwig Center for Molecular Oncology, Massachusetts Institute of Technology, Cambridge, Massachusetts

John H. Weisburger, PhD

John H. Weisburger, PhD
deceased (1921 - 2014)

Irving L. Weissman, MD

Irving L. Weissman, MD
Stanford University, Stanford, California

Zena Werb, PhD

Zena Werb, PhD
Helen Diller Family Comprehensive Cancer Center, University of California, San Francisco, San Francisco, California

Eric F. Wieschaus, PhD

Eric F. Wieschaus, PhD
Princeton University, Princeton, New Jersey

Owen N. Witte, MD

Owen N. Witte, MD
University of California, Los Angeles, Los Angeles, California

Jane Cooke Wright, MD

Jane Cooke Wright, MD
deceased (1919 – 2013)

Shinya Yamanaka, MD, PhD

Shinya Yamanaka, MD, PhD
University of California, San Francisco, San Francisco, California

Harald zur Hausen, MD

Harald zur Hausen, MD
German Cancer Research Center, Heidelberg, Germany